Justification Justification of Red List Category This species has an extremely large range, and hence does not approach the thresholds for Vulnerable under the range size criterion (extent of occurrence 30% decline over ten years or three generations). The population size is extremely large, and hence does not approach the thresholds for Vulnerable under the population size criterion (10% in ten years or three generations, or with a specified population structure). For these reasons the species is evaluated as Least Concern. Population justification In Europe, the breeding population is estimated to number 7,650,000-13,000,000 pairs, which equates to 15,300,000-26,100,000 mature individuals (BirdLife International 2015). Europe forms c.40% of the global range, so a very preliminary estimate of the global population size is 38,250,000-65,250,000 mature individuals, although further validation of this estimate is needed. Trend justification In Europe, trends between 1980 and 2013 show that populations have undergone a moderate decline (EBCC 2015).

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Ecology This species inhabits lowland and lower montane deciduous forest and woodlands, thickets and copses, as well as heaths, hedgerows and scrubby areas, parks and the edges of cultivation, including orchards and gardens in towns and cities. It occurs mostly in conifers and mixed and broadleaf forest in Siberia and other parts of Russia. In the Caucasus it favours in pine ( Pinus) and tall beech ( Fagus) forest, while in northern Europe it prefers conifer forests with well-developed undergrowth of spruce ( Picea), cedar ( Cedrus), larch ( Larix), also birch ( Betula) and yew ( Taxus). The breeding season is typically from late April to mid-September.

The nest is generally constructed of dry grass, plant fibres, roots, moss, lichen and leaves set on a loose base of dry twigs and placed up to five metres from the ground in thick bush, brambles and honeysuckle or on a flat low branch of a conifer. Clutches are four to six eggs. It feeds on a range of seeds, buds and shoots of various plants, as well as some invertebrates (Clement and Christie 2016).

The species is sedentary to migratory, with most populations likely being partially migratory (Snow and Perrins 1998).

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